Question

Where are the only two medical centres which officially store and handle specimens of the only virus known for certain to have been eradicated from the world's diseases?

Answer 1

USA and Russia
The disease is smallpox, the variola virus. In 1981, after the 33rd World Health Assembly officially declared the world free of smallpox, the four countries that either served as a WHO collaborating centre or were actively working with variola virus were the United States, England, Russia, and South Africa. By 1984, England and South Africa had either destroyed their stocks or transferred them to other approved labs. The only two locations still existing where variola virus is officially stored and handled under WHO supervision are the Centers for Disease Control and Prevention in Atlanta, Georgia, USA, and the State Research Centre of Virology and Biotechnology (VECTOR Institute) in Koltsovo, Russia.